Land and freshwater molluscs of Northern Spain

Study Extent

Locations were not predefined, but some regions were specifically visited because of known species richness or endemics occurrence. At site, observations were incidental, but microhabitats or elements that were thought to be favourable for snails or were given more attention (e.g. dead wood, north facing slopes etc.).

Sampling

Sampling was done ad random along random routes. Collection of specimens was mainly done by hand on sight. In some cases, a soil sample was taken that was later examined at home.

Quality Control

Multiple different sources were consulted for determination of the species. The dataset was reviewed by Han Raven (an expert on land and freshwater molluscs of Northern Spain) before publishing. His remarks and tips were used for improvement of the dataset.

Method steps

1. Determination was mainly done with Cadevall & Orozco (2016) and Welter-Schultes (2012) and for species with a wider, European, distribution also the following works were consulted: Cameron (2008), Glöer (2015), Horsák (2013), Jansen (2015). Finally, the following articles were consulted as well: Arconada et al. (2007) and Rolán et al. (2009) for Alzoniella, Gittenberger & Bank (1996) for Pyramidula, Gittenberger (1973) for Chondrina, Raven (1990) for Obscurella, Suárez & Quiñonero Salgado (2017) for Plentuisa vendia, Prieto et al. (2020) for Oestophora, Suárez et al. (2019), Elejalde et al. (2009) and Caro et al. (2019) for Pyrenaearia (see also Langeraert, 2017), and De Winter (1986) for Arion urbiae.

Land and freshwater molluscs do not comprise a monophyletic taxonomic group, but are all mollusc species that live in respectively land and freshwater habitats. On land, only gastropods occur (class Gastropoda, snails and slugs) and in freshwater, both gastropods as bivalves (class Bivalvia) occur.

1. Abida vasconica
2. Aegopinella nitidula
3. Alzoniella spec.
4. Ancylus fluviatilis
5. Arion (Mesarion) urbiae
6. Arion (Arion) spec.
7. Arion (Kobeltia) spec.
8. Azeca goodalli
9. Cecilioides acicula
10. Cepaea nemoralis
11. Cernuella aginnica
12. Chondrina cliedentata
13. Chondrina kobelti
14. Clausilia bidentata
15. Cochlicella acuta
16. Cochlicella barbara
17. Cochlicopa lubrica
18. Cochlicopa lubricella
19. Cornu aspersum
20. Deroceras spec.
21. Discus rotundatus
22. Elona quimperiana
23. Euglesa aff. globulare
24. Euglesa cf. casertana
25. Galba truncatula
26. Gastropoda spec.
27. Helicella itala itala
28. Helicella pampelonensis
29. Helicella spec.
30. Helicella valdeona
31. Hygromia limbata
32. Hygromiidae spec.
33. Lauria cylindracea
34. Leucophytia bidentata
35. Macrogastra rolphii digonostoma
36. Mengoana jeschaui
37. Merdigera obscura
38. Obscurella asturica
39. Obscurella asturica/hidalgoi
40. Obscurella bicostulata
41. Obscurella crassilabrum
42. Obscurella hidalgoi
43. Obscurella spec.
44. Oestophora silvae
45. Oestophorella buvinieri
46. Oligolimax annularis
47. Oxychilus navarricus navarricus
48. Paralaoma servile
49. Peringia ulvae
50. Physella acuta
51. Plentuisa vendia
52. Pomatias elegans
53. Potamopyrgus antipodarum
54. Potamopyrgus antipodarum f. carinata

Northern Spain (Vízcaya, Cantabria and Asturias)