Mecodema tenaki Seldon & Leschen 2011
- Dataset
- The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand
- Rank
- SPECIES
Classification
- kingdom
- Animalia
- phylum
- Arthropoda
- class
- Insecta
- order
- Coleoptera
- family
- Carabidae
- genus
- Mecodema
- species
- Mecodema tenaki
description
Description: Length 23.6 – 27 mm, pronotal width 6.4 – 7.4 mm, elytral width 7.4 – 8.5 mm. Colour of entire dorsal body glossy black, entire ventral body matte black. Head: Broad and flat (Fig. 14 A). Vertex smooth; vertexal groove narrow, defined entire length with V-shape medially (Fig. 56); large supraorbital puncture bearing 3 setae; 2 shallow supraorbital grooves; frons smooth with a large and very shallow depression each side of midline; frontoclypeal suture (Fig. 9) indistinct, except laterally where it meets deep tentorial pits; anterior area of clypeus with indistinct or without microsculpture; clypeus with 1 large setose puncture on each side bearing 2 – 3 setae (Fig. 56). Labrum rounded, anterior edge outwardly curved with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes squared (Fig. 12 G), median process very short and broad, slightly angled upward (15 °), strongly indentate (Fig. 12 A); setose punctures present. Submentum sclerite constriction narrow with 4 – 6 setae clustered laterally. Stipes with 2 basal setae. Gula pits large within defined suture, gula (Fig. 8) weakly convex with very fine and closely packed transverse wrinkles. Gena with well-defined isodiametric pattern across entire area. Prothorax: Prothoracic carina narrow the entire length, slightly crenulated with 4 – 6 setae each side (Fig. 56), extended beyond anterior angle; posterior lateral sinuation indistinctly carinate, slightly angled inward; pronotum broad and flattened but deflected to carina, overall shape cordate (Fig. 56); midline well-defined, fine transverse wrinkles may be present laterally on disc, small shallow dents present either side of midline; pronotal foveae deep and narrow; anterior edge slightly inwardly curved, posterior edge straight, slightly emarginated medially. Prosternum flat with fine, evenly spaced, transverse lines; proepisternum without microsculpture. Procoxal (Fig. 2) setae absent; protibia distally expanded and shovel-like (Fig. 10 B). Elytra: Broad and flattened but deflected to carina; humeral angle anteriorly convergent (Fig. 6 A); basal margin sharply angled (Fig. 56) and bevelled to base, interval 1 extended to scutellum, other intervals and striae truncated at margin; humerus with 1 – 2 setose punctures (Fig. 56); lateral carina (Fig. 11) narrow the entire length, extended to humeral angle; suture well-defined; striae with small, regularly distributed, asetose punctures; intervals weakly convex; interval microsculpture absent; 7 th strial setal pattern with 0 – 1 setose punctures in anterior ½, 3 – 4 evenly spaced setose punctures in posterior ½ (Fig. 56), setose punctures small. Ventral surface: Mesepisternum with rugose wrinkles; metepisternum without microsculpture; setose punctures present on mesocoxae (1) and metacoxae (1). All abdominal ventrites (Fig. 2) wrinkled laterally; ventrites 1 – 3 without setose punctures, ventrites 4 and 5 setose punctures present (1 – 2) each side of midline; lateral foveae absent; ventrite 6 setae present: ♂ with 2 – 4 setose punctures each side of midline, apical edge straight, ♀ with 2 – 4 setose punctures each side of midline, apical edge bluntly pointed. Anterior metaventrite process (Fig. 2) an apically broad triangle with carina poorly defined apically, narrowed posterad. Male genitalia: Apical portion of penis lobe symmetrically rounded (Fig. 56 PL) with a slight deflection to right of vertical axis (VV) (Fig. 15 C); ventral edge of penis lobe straight (VV); overall length of shaft penis lobe broad and short, slightly curved ventrally (LV) (Fig. 16 A). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; seta (scales) coverage of the apical plate moderate (26 – 75 %); size of left setose flange small; size of right setose flange large. Left paramere basal lobe ovate, narrowed slightly toward arm (Fig. 56 LP), short slope to arm; arm long and broad, narrowed to indistinct terminal lobe; apical tuft of sparse, medium-lengthed setae; setae extended along apical ¾ of ventral edge (Fig. 56 LP), ventral edge curved. Right paramere narrowly rectangular, gradually narrowed to terminus with setae extended along apical ¾ of ventral edge (Fig. 56 RP). Female genitalia: Basal gonocoxite 1 (Fig. 5) short and broad, ventral surface with a few longitudinal grooves, internal dorso-lateral carina with 2 setae. Gonocoxite 2 a short, rounded triangle with apical edge slightly reflexed. Ramus long and narrow.
description
Figure 56.
diagnosis
Diagnosis: Differs from other North Island Mecodema species by having: 1, pronotal carina with 4 - 6 setae along each side (curvidens species group); 2, pronotal carina smooth; 3, pronotal disc with a small depression each side anterad pronotal foveae; 4, shape of the basal lobe and setal distribution of the left paramere (Fig. 56 LP).
discussion
Comments: Mecodema tenaki has a very limited range, residing in degraded native kanuka (Kunzea robusta) - broadleaf forest patches in the North Cape to Cape Reinga areas.
distribution
Distribution: New Zealand, North Island, Northland, restricted to the Cape Reinga to North Cape area (see Ball et al. 2013).
materials_examined
Holotype: NZAC male labelled. NEW ZEALAND ND Cape Reinga Te Paki RBB- 7 13 Nov 2006 O. Ball / HOLOTYPE Mecodema tenaki n. sp. design. by Seldon & Leschen 2010 [red label]. Paratypes: (see Seldon & Leschen 2011).