Cryphalus Erichson 1836
- Dataset
- Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae)
- Rank
- GENUS
- Published in
- Erichson, W. F. 1836. Systematische Auseinandersetzung der Familie der Borkenkafer (Bostrichidae). Arch. Naturgeschichte 2: 45 - 65.
Classification
- kingdom
- Animalia
- phylum
- Arthropoda
- class
- Insecta
- order
- Coleoptera
- family
- Curculionidae
- genus
- Cryphalus
description
Female Eye emarginated. Antennae with three to five funicle segments. Antennal club with three visible procurved sutures, occasionally recurved, never septate. Frons convex, simple, sometimes with weak impression above epistoma, and sometimes with aciculations converging at epistoma. Pronotum with two or more marginal asperities, the margin produced in some species. Hypomeron with bifurcating setae (rare exceptions). Base of elytra sometimes with a slightly elevated ridge, straight to recurved. Angle of apex of declivity shallow, with the apex of each elytron obtuse (typically exposing part of the pygidium). Ground vestiture dense in most species (rarely sparse or absent), which can be hair-like, dagger-like or with tridentate scales. Metatibia with lateral denticles distributed over at least the apical quarter. Mesocoxae not contiguous, usually of a similar distance to distance between metacoxae or wider. Third tarsal segment weakly bilobed. Proventriculus with large apical plate, typically with sutural teeth and much larger apical teeth. Male Similar to female except sixth ventrite usually visible, frons with a transverse ridge, occasionally with enlarged setae on the protibiae and protarsi, and modified vestiture in the gular region. Penis apodemes typically as long as or longer than penis body, and separated or weakly fused at apex. Tegmen dorsally fused to form a ring. Tegmen with two short tegminal apodemes (absent in some species). Spiculum gastrale long, of a similar thickness to the penis apodemes, and without a fork. Basal sclerites not visible.
description
Where known, all species are monogamous with cave-like galleries feeding under bark.
diagnosis
Diagnosis Cryphalus can be distinguished from similar genera by the combination of emarginated eyes, sutures on the antennae clearly visible but without a septum, by the weakly bilobed third tarsal segments, the proventriculus with a large apical plate, and the aedeagus that has a tegmen sclerotized completing a ring, usually with two tegminal apodemes.
discussion
Remarks In total, 252 species known. Almost all species possess many split setae on the hypomeron which can easily separate species in this genus from most similar bark beetles. The presence, number, and arrangement of sutural teeth on the apical plate of the proventriculus have been extensively used to diagnose Chinese species (see Tsai and Li (1963 )). Many species are described. It is likely that several of the species names should be synonymized, but many types are in a poor condition with few external diagnostic characters known. The genus was previously split into three genera, including Hypocryphalus and Margadillius based primarily on the number of funicular segments. This is, however, a rapidly evolving character which may vary within a species and has a poor phylogenetic signal. The large number of species, particularly those described by Schedl, create many secondary homonyms. Where appropriate, these have been corrected.
distribution
Distribution Africa, Asia, Europe, Oceania, North America, Central America (introduced), South America (introduced).
materials_examined
Type material examined Allotype of Ernoporus antennarius Schedl, 1974 (ANIC); paratype of Ernoporus antennarius Schedl, 1974 (NHMW); holotype of Taenioglyptes aquilonius Nobuchi, 1975 (ITLJ); holotype of Cryphalus araucariae Schedl, 1970 (ANIC); holotype of Taenioglyptes artestriatus Browne, 1970 (BMNH); holotype of Ericryphalus artocarpus Schedl, 1939 (BMNH); holotype and paratype of Cryphalus asperulus Schedl, 1948 (BMNH); paratype of Cryphalus asperulus Schedl, 1948 (BMNH); holotype of Cryphalus ater Browne, 1984 (BMNH); holotype of Cryphalus basihirtus Beeson, 1929 (BMNH); holotype of Taenioglyptes bicarinatus Nobuchi, 1975 (ITLJ); holotype of Cryphalomorphus bicolor Browne, 1983 (BMNH); holotype of Hypocryphalus bidentatus Browne, 1980 (BMNH); holotype of Cryphalus brasiliensis Schedl, 1976 (NHMW); paratype of Ptilopodius brevis Browne, 1970 (BMNH); allotype of Cryphalus brevisetosus Schedl, 1943 (NHMW); lectotype of Cryphalus brevisetosus Schedl, 1943 (NHMW); holotype of Cryphalus brimblecombei Schedl, 1948 (BMNH); holotype of Cryphalus brunneus Browne, 1984 (BMNH); holotype of Cryphalus buloloensis Browne, 1984 (BMNH); paratype of Cryphalus buloloensis Browne, 1984 (BMNH); holotype of Margadillius carinatus Browne, 1980 (BMNH); paratype of Cryphalus carpini Berger, 1917 (ZIN); holotype of Cryphalus cylindricus Browne, 1980 (BMNH); holotype of Cryphalus cylindrus Browne, 1984 (BMNH); holotype of Dacryphalus cylindrus Browne, 1950 (BMNH); paratype of Cryphalus cylindrus Browne, 1984 (BMNH); holotype of Cryphalus dilutus Eichhoff, 1878 (NHMW); holotype of Cryphalus dipterocarpi Wood, 1989 (USNM); holotype of Taenioglyptes dissimilis Nobuchi, 1975 (ITLJ); holotype and paratype of Cryphalus diversicolor Browne, 1984 (BMNH); holotype of Cryphalus exiguus Blandford, 1894 (BMNH); holotype of Euptilius exiguus Browne, 1984 (BMNH); holotype of Cryphalus felis Wood, 1989 (USNM); holotype of Hypocryphalus fici Browne, 1986 (BMNH); holotype of Hypocryphalus froggatti Nunberg, 1961 (BMNH); holotype of Cryphalus fuliginosus Blandford, 1895 (BMNH); holotype of Cryphalus fulmineus Wood, 1989 (USNM); holotype of Margadillius fulvus Browne, 1984 (BMNH); cotype of Cryphalus garciniae Nobuchi, 1959 (ITLJ); holotype of Cryphalus grayi Schedl, 1968 (ANIC); holotype of Hypothenemus griseus Blackburn, 1885 (BMNH); holotype of Cryphalus helopioides Schedl, 1953 (BMNH); holotype of Taenioglyptes hirsutus Nobuchi, 1975 (ITLJ); holotype of Cryphalus inops Eichhoff, 1872 (RBINS); holotype of Taenioglyptes kagoshimensis Nobuchi, 1975 (ITLJ); holotype and paratype of Cryphalus kesiyae Browne, 1975 (BMNH); paralectotype of Cryphalus kurenzovi Stark, 1936 (ZIN); holotype of Hypocryphalus laevis Browne, 1980 (BMNH); holotype and paratype of Cryphalus laevis Browne, 1984 (BMNH); type of Cryphalus laricis Niisima, 1909 (ITLJ); holotype and paratype of Hypocryphalus laticollis Browne, 1974 (BMNH); holotype of Cryphalus longior Browne, 1984 (BMNH); paratype of Cryphalus longior Browne, 1984 (BMNH); holotype and two paratypes of Taenioglyptes longipennis Browne, 1970 (BMNH); holotype of Hypocryphalus longipilis Browne, 1981 (BMNH); holotype of Taenioglyptes longisetosus Nobuchi, 1975 (ITLJ); holotype and paratype of Margadillius magnus Browne, 1984 (BMNH); holotype of Hypocryphalus malayensis Schedl, 1942 (BMNH); syntype of Cryphalus mandschuricus Eggers, 1929 (ITLJ); paralectotype and lectotype of Cryphalus mangiferae Stebbing, 1914 (BMNH); holotype of Margadillius margadilaonis Hopkins, 1915 (USNM); holotype of Taenioglyptes meridionalis Nobuchi, 1975 (ITLJ); Two paralectotypes of Cryphalus minimus Eggers, 1927 (NHMW, BMNH); holotype of Hypocryphalus minutus Browne, 1980 (BMNH); holotype of Cryphalus mollis Schedl, 1955 (BMNH); holotype of Cryphalus montanus Nobuchi, 1964 (ITLJ); paratype of Hypocryphalus montanus Schedl, 1974 (NHMW); holotype and paratype of Cryphalus negrosensis Browne, 1979 (BMNH); holotype of Cryphalus nitens Browne, 1980 (BMNH); holotype and paratype of Cryphalus nitidipennis Browne, 1984 (BMNH); paratype of Cryphalus nothofagi Browne, 1984 (BMNH); holotype of Margadillius papuanus Schedl, 1973 (ANIC); paratype of Margadillius papuanus Schedl, 1973 (NHMW); lectotype of Cryphalus perminimus Schedl, 1942 (NHMW; holotype of Cryphalus pilosulus Browne, 1980 (BMNH); holotype of Taenioglyptes pulchellus Nobuchi, 1975 (ITLJ); holotype and paratype of Cryphalus punctatus Browne, 1984 (BMNH); holotype of Margadillius quadrituberculatus Schedl, 1963 (NHMW); holotype of Hypocryphalus reflexus Browne, 1980 (BMNH); syntypes of Cryphalus rhusii Niisima, 1909 (ITLJ); syntypes (6) of Cryphalus robustus Eichhoff, 1872 (RBINS (5), BMNH (1 )); holotype of Hypocryphalus rotundus Hopkins, 1915 (USNM); holotype of Ericryphalus rugosus Schedl, 1958 (BMNH); holotype of Cryphalus samoensis Beeson, 1929 (BMNH); syntype of Cryphalus sandakanensis Schedl, 1937 (BMNH); holotype of Cryphalus sawadai Nobuchi and Takahashi, 1965 (ITLJ); holotype of Cryphalus scabripennis Schedl, 1972 (BMNH); holotype of Ericryphalus securus Schedl, 1940 (BMNH); holotype of Taenioglyptes sordidus Nobuchi, 1975 (ITLJ); holotype of Cryphalomorphus striatulus Browne, 1978 (BMNH); holotype of Cryphalus striatulus Browne, 1981 (BMNH); holotype of Cryphalus striatus Browne, 1974 (BMNH); lectotype and 3 paralectotypes Cryphalus swezeyi Schedl, 1942 (BMNH (2), NHMW (2 )); syntype of Cryphalus swezeyi Schedl, 1942 (BMNH); holotype of Hypothenemus sylvicola Perkins, 1900 (BMNH); holotype of Cryphalus terminaliae Browne, 1980 (BMNH); holotype of Margadillius terminaliae Browne, 1984 (BMNH); syntype of Cryphalus trypanus Sampson, 1914 (BMNH); holotype of Cryphalus vestitus Blandford, 1895 (BMNH); holotype and paratype of Cryphalus vitiensis Browne, 1974 (BMNH); holotype of Cryptarthrum walkeri Blandford, 1896 (BMNH).
type_taxon
Type of genus Bostrichus asperatus Gyllenhal, 1813.
Name
- Synonyms
- Acryphalus Tsai and Li 1963
- Homonyms
- Cryphalus Erichson 1836