Haptoderus colasi tuberosus J.P.Nicolas, 1973 Dataset GBIF Backbone Taxonomy Rank SUBSPECIES Classification kingdom Animalia phylum Arthropoda class Insecta order Coleoptera family Carabidae genus Pterostichus species Pterostichus colasi Name Homonyms Haptoderus colasi tuberosus J.P.Nicolas, 1973