Patrera procera (Keyserling 1891)

Brescovit, 1997: 50. Patrera procera: Brescovit, 1997: 33, 50.

Description. Male (Lectotype, BMNH). Carapace orange brown, with orange paramedian stripes. Chelicerae brown yellowish, labium and endites orange. Sternum light orange, with dark borders. Legs orange. Abdomen dorsally brownish, ventrally cream (Fig. 1 A). Total length 8.6, carapace length 3.5, width 2.65. Clypeus height 0.15. Eye diameters and interdistances: AME 0.11, ALE 0.20, PME 0.17, PLE 0.20; AME – AME 0.06, AME – ALE 0.08, PME – PME 0.21, PME – PLE 0.17, ALE – PLE 0.07. Chelicerae 2.3 long, four promarginal teeth, nine small retromarginal denticles (Fig. 3 A). Leg measurements: leg I — femur 5.5 / patella 1.5 / tibia 6.1 / metatarsus 5.2 / tarsus 2.05 / total 20.35; II — 4.85 / 1.3 / 5.25 / 4.6 / 1.9 / 17.9; III — 3.1 / 1.05 / 2.5 / 3.4 / 1.0 / 11.05; IV — 4.8 / 1.3 / 4.35 / 5.75 / 1.2 / 17.75. Leg spination: I — II — tibia v 2 - 2 - 0, p 1 - 1 - 1, r 1 - 1 - 1, metatarsus v 2 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0; III — IV — tibia v 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, metatarsus v 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1. Abdomen: length 5.35, epigastric furrow 1.4 from tracheal spiracle, spiracle 2.3 from base of spinnerets. Palp: tibia as long as cymbium; tegulum oval; spermatic duct “ Ushaped ” ventrally; short distal tegular projection, close to the embolus apex; subdistal median apophysis, inverted “ S-shaped ”; embolus with median prolateral origin, thin, with broad base, slightly curved (Figs 1 C, 2 A, 3 B, C). Female (Paralectotype BMNH). Coloration as in male, except dorsal area of the abdomen lighter (Fig. 1 B). Total length 9.2, carapace length 3.8, width 2.8. Clypeus height 0.16. Eye diameters and interdistances: AME 0.11, ALE 0.20, PME 0.16, PLE 0.18; AME – AME 0.07, AME – ALE 0.06, PME – PME 0.22, PME – PLE 0.18, ALE – PLE 0.06. Chelicerae 1.67 long, five promarginal teeth, eight retromarginal denticles. Leg measurements: leg I — femur 4.8 / patella 1.55 / tibia 5.1 / metatarsus 4.1 / tarsus 2.1 / total 17.65; II — 4.3 / 1.4 / 4.5 / 3.85 / 1.7 / 15.75; III — 3.1 / 1.2 / 2.4 / 3.1 / 1.0 / 10.8; IV — 4.6 / 1.4 / 4.1 / 5.0 / 5.0 / 1.3 / 16.4. Leg spination: I — tibia v 2 - 2 - 0, p 1 - 0 - 0, r 1 - 0 - 0, metatarsus v 2 - 0 - 0, p 0, r 0; II — tibia v 2 - 2 - 0, p 1 - 1 - 1, r 0, metatarsus v 2 - 0 - 0, p 1 - 0 - 0, r 0; III — IV — tibia v 2 - 2 - 2, p 1 - 1 - 1 -, r 1 - 1 - 1, metatarsos v 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1. Abdomen: length 5.7, epigastric furrow 1.55 from tracheal spiracle, spiracle 2.25 from base of spinnerets. Epigynum: lateral borders narrow, not distally joined; internally with long, coiled copulatory ducts; seminal receptacles in the proximal area of the copulatory ducts; oval spermathecae; fertilization ducts as long as the spermathecae, curved and basally inserted (Figs 1 F, G, 2 C, D, 3 F). Variation. Males (n = 10): total length 7.4 – 9.2; carapace length 2.8 – 3.8; femur I length 3.8 – 5.2. Females (n = 10): total length: 8.5 – 9.7; carapace length 3.3 – 3.6; femur I length 4.2 – 4.8.

Figs 1 A – G; 2 A – D; 3 A – F; 27 C

Diagnosis. Males of Patrera procera can be distinguished from the remaining species by the characteristic tibial retrolateral apophysis wide, subrectangular, with a transverse fold at the tip, forming a conspicuous groove, and by the conical dorsal tibial apophysis (Figs 1 D, 2 B, 3 E) in the male palp. Females are diagnosed by their short, narrow epigynal lateral borders with lateral excavation at the internal borders and wide deep atrium in the epigynum (Figs 2 C, 3 F).

Distribution. Northeast to southern Brazil, West Paraguay and Northeastern Argentina (Fig. 27 C).

Other material examined. BRAZIL. No date, 1 ♂ (MNHN 18666); Paraíba: João Pessoa, Mata do Buraquinho (7 ° 8 ′ 17 ″ S; 34 ° 50 ′ 42 ″ W), 1 ♀, 2003 – 2004, S. C. Dias leg. (IBSP 99968); Areia, Reserva da Mata do Pau Ferro (06 º 57 ’ S; 35 º 44 ’ W), 1 ♂, 23 – 29. IX. 1999, A. D. Brescovit et al. leg. (IBSP 138929); 2 ♂ (IBSP 138939); 1 ♂ (IBSP 138949); 1 ♂ (IBSP 138950); 1 ♂ (IBSP 138958); 1 ♂ (IBSP 138960); all collected in 23 – 29. IX. 1999 by A. D. Bresco- vit et al.; Bahia: 2 ♂ (MNHN 7135); Una, Reserva Biológica do Una (15 ° 10 ′ 23 ″ S; 39 ° 7 ′ 56 ″ W), 1 ♀, 15 – 28. XI. 2000, A. D. Brescovit et al. leg. (IBSP 45700); Jussari (15 ° 11 ’ 27 ” S; 39 ° 29 ’ 42 ” W), 3 ♀, 1970, Cepec / Ceplac leg. (MNRJ 3143); 1 ♂, 1 ♀ (MNRJ 3236); 1 ♂ (MNRJ 3194); 2 ♂ (MNRJ 3256); 1 ♂, 2 ♀ (MNRJ 13313); 2 ♂ (MNRJ 3183); Coaraci (14 ° 38 ′ 27 ″ S; 39 ° 33 ′ 3 ″ W), Fazenda Boa Esperança, 1 ♂, 01. X. 1970, Cepec / Ceplac leg. (MNRJ 3231); Mato Grosso do Sul: Bonito, Baia Bonita (21 º 09 ’ S; 56 º 26 ’ W), 1 ♂ (IBSP 138887); 1 ♀ (IBSP 138890); 3 ♂, 2 ♀ (IBSP 138902); all collected in 14 – 23. X. 2002, Equipe Biota; Minas Gerais: Belo Horizonte, Estação Ecológica UFMG, Malaise trap (19 ° 52 ′ 19 ″ S 43 ° 57 ′ 58 ″ W), 2 ♂, 09 – 16. XI. 1999, J. C. R. Fontenelle leg. (IBSP 41119); Espírito Santo: São José do Calçado, Rio Itabapoana, Usina Hidrelétrica de Rosal (20 ° 57 ’ 7 ” S; 41 ° 43 ’ 5 ” W), 1 ♂, 1 ♀, XI. 1999, I. Knysak leg. (IBSP 26383); Apiacá (21 ° 9 ′ 14 ″ S; 41 ° 34 ′ 4 ″ W), 1 ♂, XI. 1986, R. L. C. Baptista leg. (MNRJ 3188); Ca- riacica, Alto do Alegre, Reserva Estadual de Duas Caras (20 ° 17 ’ 29.7 ” S; 40 ° 31 ’ 10.2 ” W), 1 ♂, 13. X. 2005, T. Souza et al. leg. (IBSP 133182); Linhares, Flona de Goytacazes (19 ° 23 ’ 48 ” S; 40 ° 3 ’ 42 ” W), 42 m alt., 2 ♂, 21 – 23. X. 2005, T. Souza et al. leg. (IBSP 134825); Conceição da Barra, Floresta Nacional do Rio Preto, Trilha do Mata-Pau, 60 m alt. (18 ° 24 ’ 00 ” S; 39 ° 50 ’ 00 ” W), 1 ♂, 19. X. 2005, T. Souza et al. leg. (IBSP 135120); Rio de Janeiro: Volta Redonda, Floresta da Cicuta (22 ° 32 ′ 53 ″ S; 44 ° 5 ′ 34 ″ W), 3 ♀, VIII. 2002, E. L. C. Silva et al. leg. (IBSP 48942; IBSP 48950; IBSP 48954); Cachoeiras de Macacu, Reserva Ecológica de Guapiaçu (22 ° 35 ′ 14.61 ″ S; 42 ° 56 ′ 24.69 ″ W), 11 ♂ (IBSP 138968; IBSP 138970; IBSP 138972; IBSP 138974; IBSP 138984; IBSP 138991; IBSP 138993; IBSP 138994; IBSP 138997); all collected in 08 – 12. X. 2001 by Equipe Biota; Pinheiral, Fazenda Santa Helena (22 º 34 ’ 00 ” S; 44 º 21 ’ 00 ” W), 21 ♂, 3 ♀ (IBSP 139401; IBSP 139406 - IBSP 139408; IBSP 139410; IBSP 139411; IBSP 139414; IBSP 139415; IBSP 139418 - IBSP 139420; IBSP 139423 - IBSP 139427; IBSP 139430; IBSP 139432 - IBSP 139434); all collected in 05 – 11. X. 1999 by A. D. Brescovit et al.; Pinheiro (currently Piraí, Pinheiral) (22 ° 30 ′ 46 ″ S; 44 ° 00 ′ 3 ″ W), 2 ♂, I. 1988, R. L. C. Baptista leg. (MNRJ 3189); 1 ♂, 1 ♀, 4 # imm. [without date], C. F. de Mello-Leitão leg. (MNRJ 616); São Paulo: Teodoro Sampaio, Porto Cabral, Rio Paraná (24 ° 31 ’ 00.1 ” S; 47 ° 42 ’ 00 ” W), 1 ♀, 1941, L. P. Travassos Filho leg. (MZSP 9788); Ribeirão Preto (21 ° 10 ′ 40 ″ S; 47 ° 48 ′ 36 ″ W), 1 ♂, 1992, E. Camillo leg. (IBSP 210237); Itapetin- inga, Estação Ecológica de Angatuba (23 º 24 ’ 00 ” S; 48 º 21 ’ 00 ” W), 2 ♂, 7 ♀ (IBSP 138710; IBSP 138711; IBSP 138714; IBSP 138717; IBSP 138720; IBSP 138722; IBSP 138723; IBSP 138725), all collected in 11 – 16. XI. 2002 by Equipe Biota; Iporanga (24 ° 35 ’ 09 ” S; 48 ° 35 ’ 34 ” W), Lajeado, 1 ♂, XII. 1944, J. Damico leg. (MZSP 12217); Itaberá (23 ° 51 ′ 43 ″ S; 49 ° 8 ′ 13 ″ W), 2 ♂, 1 ♀, XI. 1943, F. S. Pereira leg. (MZSP 12207); Botucatu, Fazenda Butignoli, Rubião Júnior (23 ° 59 ’ 21 ” S; 48 ° 26 ’ 21 ” W), 1 ♀, 07. I. 1987, I. M. P. Rinaldi & L. C. Forti leg. (MCN 22486); 1 ♂, 07. X. 1987, I. M. P. Rinaldi & L. C. Forti leg. (MCN 22485); 1 ♀, 12. II. 1987, I. M. P. Rinaldi leg. (MZSP 60386); 2 ♀, 18. XII. 1978, N. Carneiro leg. (MZSP 12129); 1 ♀, 02. XII. 1987, I. M. P. Rinaldi & L. C. Forti leg. (MCN 22483); 1 ♂, 02. XII. 1987, I. M. P. Rinaldi & L. C. Forti leg. (MCN 22484); 3 ♂, 1 ♀, 14 – 16. XII. 1966, (MZSP 12123; MZSP 12141; MZSP 12317; MZSP 14137); all collected by V. C. de Jesus; 1 ♂, 16. XII. 1986, I. Rinaldi leg. (MZSP 60360); Botucatu (22 ° 53 ’ 9 ” S; 48 ° 26 ’ 42 ” W), Parque Municipal, 1 ♀, 11. V. 1986 (MZSP 60383); 1 ♂, 12. II. 1987 (MZSP 60385); 1 ♂, 13. I. 1988 (MZSP 60361); 3 ♀, 02. VIII. 1988 (MZSP 60362); 3 ♀, 12. II. 1987 (MZSP 60363), all collected by I. Rin- aldi; Fazenda Edgárdia, campus UNESP (22 ° 48 ’ 56 ” S; 48 ° 23 ’ 44 ” W), 1 ♀, 07. VI. 1988, I. M. P; Rinaldi leg. (MZSP 60384); Rio Claro (22 ° 24 ′ 39 ″ S; 47 ° 33 ′ 39 ″ W), 1 ♀, 17 – 19. I. 1962, H. R. Filho leg. (MZSP 10216); Assis, Estação Ecológica de Assis (22 º 34 ’ 00 ” S; 50 º 24 ’ 00 ” W), 1 ♂, 3 ♀, 25 – 30. XI. 2002, Equipe Biota leg. (IBSP 138827; IBSP 138842; IBSP 138860); Atibaia (23 º 07 ’ 01 ’’ S; 46 º 33 ’ 01 ” W), 1 ♂, 13. X. 2002, G. Machado et al. leg. (IBSP 35177); 1 ♂, X. 2002, M. B. Rosseto & G. Machado leg. (IBSP 35845); Biritiba Mirim (23 ° 34 ′ 22 ″ S; 46 ° 2 ′ 20 ″ W), 1 ♂, 2003, I. Knysak leg. (IBSP 72718); Jundiaí (23 º 11 ’ 11 ” S; 46 º 53 ’ 03 ” W), 1 ♂, 09 – 12. X. 1999, C. A. Rheims leg. (IBSP 26252); Reserva Biológica Municipal da Serra do Japi (23 ° 12 ’ 00 ” S; 46 º 30 ’ 00 ” W), 2 ♂, 2004, Alunos Curso Pós USP leg. (IBSP 138871; IBSP 138872); 1 ♀, III. 2007, A. D. Brescovit et al. leg. (IBSP 215510); 1 ♂, 2 ♀, 15 – 17. XII. 2014, Exc. Alunos USP leg. (IBSP 231027); Cabreúva (23 º 18 ’ 27 ” S; 47 º 07 ’ 58 ’’ W), 27. X. 2001, D. F. Candi- ani & P. E. D. Francesconi leg. (IBSP 31976); Itapecerica da Serra (23 ° 43 ′ 1 ″ S; 46 ° 50 ′ 56 ″ W), 1 ♂, 10. I. 2006, L. V. de Camões leg. (IBSP 75579); Mogi das Cruzes, Rio Tietê (23 ° 31 ′ 22 ″ S; 46 ° 11 ′ 16 ″ W), 1 ♀, VII – VIII. 1996, R. Martins leg. (IBSP 13831); Barueri (23 ° 30 ′ 38 ″ S; 46 ° 54 ′ 34 ″ W), 1 ♀, 07. I. 1961 (MZSP 12250); 2 ♀, 31. XII. 1965 (MZSP 14108); 2 ♀, 21. XII. 1965 (MZSP 14139), 1 ♂, 08. I. 1966 (MZSP 5958); 1 ♂, 08. I. 1966 (MZSP 12314); 3 ♂, 1 ♀, 31. XI. 1965 (MZSP 14136); 1 ♂, 1 ♀, 05. II. 1965 (MZSP 14138); all collected by K. Lenko; Embu das Artes (23 ° 38 ′ 56 ″ S; 46 ° 51 ′ 7 ″ W), 1 ♀, II. 1946, F. Lane leg. (MZSP 12242); São Paulo (23 º 32 ’ 51 ” S; 46 º 38 ’ 10 ” W), 1 ♀, X. 1990, Equipe IBSP leg. (IBSP 4367); Vila Mascote (23 ° 38 ’ 44 ” S; 46 ° 40 ’ 2 ” W), 1 ♀, 04. XI. 2002, D. M. B. Battesti leg. (IBSP 36392); Perdizes (19 ° 21 ’ 19 ’’ S; 47 ° 16 ’ 58 ’’ W), 1 ♂, 01. XII. 1951, H. Urban leg. (MZSP 12276); Parque da Previdência (23 º 34 ’ 00 ” S; 45 º 43 ’ 00 ” W), 1 ♂, 22. X. 1999, F. S. Cunha leg. (IBSP 29400); 1 ♂, 29. X. 1999, J. Im- pério leg. (IBSP 29401); Campus USP, Reserva Florestal da CUASO (23 º 33 ’ 00 ” S; 46 º 43 ’ 00 ” W), 3 ♀, 14. XII. 1999 (IBSP 29393); 1 ♀, 29. X. 1999 (IBSP 29394); 1 ♂, 29. X. 1999 (IBSP 29395); 1 ♀, 31. X. 2000 (IBSP 29396); 1 ♀, 14. XII. 1999 (IBSP 29398); 1 ♂, 27 / X / 1999 (IBSP 32984); 2 ♀, all collected by F. S. Cunha; 29. X. 1999, D. F. Candi- ani leg. (IBSP 29399); 1 ♂, 05. IX. 2000, J. Império leg. (IBSP 29397); Campus Instituto Butantan (23 º 33 ’ 00 ” S; 46 º 43 ’ 00 ” W), 4 ♂, 2 ♀, 11. XI. 2004, no coll. (IBSP 48427); 2 ♀, 2010, G. S. Ruiz leg. (IBSP 168580); Campus Insti- tuto Butantan, Mata IBSP, 5 ♂, 6 ♀, 05. X. 2005, Alunos USP curso 2006 leg. (IBSP 57444); Represa de Guarapir- anga, Ilha dos Eucaliptos (23 ° 44 ’ 26 ” S; 46 ° 44 ’ 09 ” W), 1 ♂, 09. IX. 2001, R. P. Indicatti leg. (IBSP 32580); Ilha dos Eucaliptos, 4 ♂ 8 ♀ (IBSP 61264; IBSP 61265; IBSP 61266; IBSP 61267; IBSP 61268; IBSP 61269; IBSP 61270; IBSP 61271; IBSP 61272); all collected in 07 – 13. X. 2004 by I. Cizauskas & C. R. M Garcia; Parque Estadual do Jaraguá (23 ° 27 ′ 34.3 ″ S; 46 ° 46 ′ 2.8 ″ W), 2 ♂, 2 ♀, 27 – 28. X. 2005, Curso UFRRJ leg. (IBSP 71185; IBSP 71237; IBSP 71239); 10 ♂, 4 ♀, 14 – 19. X. 2002, Equipe Biota leg. (IBSP 139596; IBSP 139598; IBSP 139599; IBSP 139602; IBSP 139603; IBSP 139605; IBSP 139615; IBSP 139616; IBSP 139618; IBSP 139624; IBSP 139636; IBSP 139647; IBSP 139648); Parque Estadual da Cantareira, Sede (23 ° 24 ′ 5 ″ S; 46 ° 35 ′ 24 ″ W), 1 ♀, 20. XII. 2000 (IBSP 85701); 2 ♀, 21. XII. 2000 (IBSP 85702); 1 ♀, 28. II. 2001 (IBSP 85703); 2 ♂, 15. XII. 2000 (MCN 41880; MCN 41882); 2 ♂, 3 ♀ (MZSP 24966 – MZSP 24970), all collected in 22. XII. 2000 by R. Pinto da Rocha et al.; Núcleo Pedra Grande, 1 ♀, 15. XII. 2002, R. Pinto da Rocha et al. leg. (IBSP 210221); 3 ♂, 3 ♀ (MZSP 24959 – MZSP 24964), all collected in 21. XII. 2000 - 6. I. 2001 by R. Pinto da Rocha et al.; Parque Estadual Fontes do Ipiranga (23 ° 38 ′ 24.6 ″ S; 46 ° 37 ′ 3.1 ″ W), 1 ♀, 28. X. 2017 (IBSP 222758); 1 ♂, 27. VII. 2018 (IBSP 222766); 1 ♀, 28. X. 2017 (IBSP 222768); 1 ♀, 25. XI. 2017 (IBSP 222769); 1 ♂, 30. IX. 2017 (IBSP 222777); 1 ♂, 27. VIII. 2017 (IBSP 222778); 1 ♂, 22. VII. 2017 (IBSP 222780); 1 ♀, 25. XI. 2017 (IBSP 222783); 1 ♂, 30. IX. 2017 (IBSP 222787); 1 ♂, 27. VIII. 2017 (IBSP 222792); 1 ♀, 27. VIII. 2017 (IBSP 222793); 1 ♀, 27. VIII. 2017 (IBSP 222794), all collected by Z. R. Mendes; Horto Florestal (currently Parque Estadual Alberto Löfgren) (23 ° 27 ′ 33 ″ S; 46 ° 38 ′ 2 ″ W), 1 ♂, 1 imm., 1. XI. 1994, R. Pinto da Rocha leg. (MZSP 14310); Bosque da Saúde (23 ° 34 ’ 21 ” S; 46 ° 38 ’ 70 ” W), 2 ♀, 09. IV. 1944, F. Lane leg. (MZSP 5165); 1 ♂, 22. III. 1942, F. Lane leg. (MZSP 14134); Cocaia, Represa Billings (23 ° 47 ’ 13 ” S; 46 ° 35 ’ 2 ” W), 1 ♀, 11. III. 1951, H. Urban leg. (MZSP 12212); Parque Estadual Serra da Cantareira, Sede (23 ° 24 ′ 5 ″ S; 46 ° 35 ′ 24 ″ W), 1 ♂, 22. XI. 2000, R. Pinto da Rocha et al. leg. (MZSP 25011); São José do Barreiro, Parque Nacional da Serra da Bocaina (23 ° 2 ′ 30 ″ S; 44 ° 39 ′ 42 ″ W), 1 ♂, 22 – 25. VII. 2006, F. U. Yamamoto leg. (IBSP 66747); Ribeirão Grande, Parque Estadual de Intervales (24 ° 16 ′ 6.3 ″ S; 48 ° 24 ′ 49.7 ″ W), in Bamboo, 1 ♀, 2002 – 2003, M. O. Gonzaga leg. (IBSP 71452); São Bernardo do Campo (23 ° 41 ′ 38 ″ S; 46 ° 33 ′ 54 ″ W), Clube de Campo São Camilo, 1 ♂, 1 ♀, 20. X. 2008, V. Passanha & C. V. Janini leg. (IBSP 119790); Mairiporã (23 ° 22 ′ 44 ″ S; 46 ° 31 ′ 38 ″ W), Serra da Cantareira, 1 ♂, VII. 2000, C. R. Macagnan leg. (IBSP 27347); Mogi Guaçu, Reserva Biológica Mogi Guaçu (22 º 10 ’ 00 ” S; 47 º 08 ’ 00 ” W), 2 ♀, 16. XII. 1998, E. Camillo leg. (IBSP 28113; IBSP 28155); Reserva Biológica Mogi Guaçu (22 º 10 ’ 00 ” S; 47 º 08 ’ 00 ” W), 33 ♂, 2 ♀ (IBSP 54646; IBSP 139460; IBSP 139466; IBSP 139468; IBSP 139469; IBSP 139473; IBSP 139482; IBSP 139484; IBSP 139486; IBSP 139489; IBSP 139491; IBSP 139493; IBSP 139497; IBSP 139500; IBSP 139501; IBSP 139502; IBSP 139504; IBSP 139505; IBSP 139509; IBSP 139510; IBSP 139511; IBSP 139512; IBSP 139514; IBSP 139516; IBSP 139517; IBSP 139518), all collected in 13 – 19 / X / 2003 by Equipe Biota; Pariquera-Açu (24 ° 42 ’ 54 ” S; 47 ° 52 ’ 51 ” W), Vale do Ribeira, cam- pus do Instituto Agronômico de Campinas, 1 ♂, [without date], E. Wienskoski leg. (MZSP 60391); Ubatuba (23 º 26 ’ 02 ” S; 45 º 04 ’ 16 ” W), Fazenda Experimental, 1 ♂, 1 ♀, 14. XI. 1943, A. Zoppei leg. (MZSP 12228); Paraná: Cruzeiro do Iguaçu, Usina Foz do Chopim (25 º 33 ’ 45 ” S; 53 º 10 ’ 37 ” W), 1 ♂, 08 – 15. X. 1998, Equipe IBSP leg. (IBSP 21241); Adrianópolis, Vale do Ribeira (24 ° 39 ′ 25 ″ S; 48 ° 59 ′ 27 ″ W), 1 ♀, 18. XI. 2004, D. M. B. Battesti leg. (IBSP 47949); Bela Vista do Paraíso (22 º 59 ’ 49 ” S; 51 º 11 ’ 27 ” W), 1 ♂, 1 ♀, 11 – 16. XI. 1991, A. B. Bonaldo leg. (IBSP 221089); Fênix (23 º 54 ’ 57 ” S; 51 º 58 ’ 44 ” W), 1 ♂, 24. XI. 1986, Equipe CIIF leg. (IBSP 221097); 1 ♂, 01. XI. 1986, Equipe Profaupar leg. (IBSP 221098); Parque Estadual de Vila Rica do Espírito Santo (23 ° 55 ’ 00 ” S; 51 ° 57 ’ 00 ” W), 3 ♀, 12. XI. 1987, A. D. Brescovit leg. (IBSP 221096); Curitiba, campus UFPR, Centro Politécnico (25 ° 25 ′ 44 ″ S; 49 ° 16 ′ 3 ″ W), 2 ♂, 2 ♀, 26. X. 1995, A. B. Bonaldo leg. (MCN 26698); Almirante Tamandaré (25 ° 19 ′ 30 ″ S; 49 ° 18 ′ 36 ″ W), 1 ♀, 10. VII. 1984 (MCN 12389); 1 ♂, 25. IV. 1984 (MCN 12414); 1 ♂, 08. VI. 1984 (MCN 12454); 1 ♂, 25. IV. 1984 (MCN 12467); 1 ♂, 22. VI. 1984 (MCN 12488); 1 ♂, 28. IX. 1984 (MCN 12517); 1 ♂, 28. IX. 1984 (MCN 12518); all collected by E. C. Costa; Capanema (25 ° 40 ′ 19 ″ S; 53 ° 48 ′ 32 ″ W), 3 ♂, 11. III. 1981, S. Scherer leg. (MCN 9777); Santa Catarina: Chapecó, Floresta Nacional de Chapecó (27 º 06 ’ 10,5 ” S; 52 º 46 ’ 48,8 ” W), 1 ♀, I. 2003 – XII. 2004, M. Scartezini leg. (IBSP 91070); Nova Teutônia (27 ° 11 ’ 00 ” S; 52 ° 23 ’ 00 ” W), 2 ♂, 1 ♀, [without date], F. Plaumann leg. (FMS 13579; FMS 13583; FMS 13578); Florianópolis, Itacorubí, UCAD (27 ° 33 ′ 21.44 ″ S; 48 ° 29 ′ 37.46 ″ W), 1 ♂, 2004, J. Steiner leg. (IBSP 91097); Criciúma (28 ° 40 ′ 40 ″ S; 49 ° 22 ′ 12 ″ W), 1 ♂, 08. XI. 2007, R. A. Teixeira leg. (IBSP 132769); Rancho Queimado (27 ° 40 ′ 22 ″ S; 49 ° 1 ′ 19 ″ W), 1 ♂, 15 – 18. XI. 1995, L. A. Moura leg. (MCN 27039); 2 ♀, 13 – 15. I. 1995, A. B. Bonaldo leg. (MCN 26356); Rio Grande do Sul: Iraí (27 ° 11 ′ 38 ″ S; 53 ° 15 ′ 3 ″ W), 1 ♂, 2 ♀, 20. XI. 1975, A. A. Lise leg. (MCN 3106); Derrubadas, Parque Estadual do Turvo, Salto do Yucumã (27 ° 08 ’ 32.5 ″ S; 53 ° 53 ’ 09.0 ″ W), 1 ♀, 16. I. 1985 (MCN 13002); 1 ♂, 16. I. 1985 (MCN 13005); all collected by A. A. Lise; São Fran- cisco de Paula (29 ° 26 ′ 52 ″ S; 50 ° 35 ′ 2 ″ W), 1 ♀, 22. IX. 1974 (MCN 2304); 1 ♂, 1 ♀, 22. I. 1981 (MCN 9531); 1 ♂, 2 ♀, 23. I. 1981 (MCN 9540); all collected by A. A. Lise; Barragem dos Bugres (29 ° 20 ’ 35.87 ” S; 50 ° 41 ’ 44.15 ” W), 2 ♂, 23 – 25. XI. 1998, A. F. Franceschini leg. (MCN 30656); 2 ♂, 1 ♀, 01 – 04. II. 1999, L. A. Moura leg. (MCN 30669); Rincão dos Kroeff (29 ° 28 ’ 3 ” S; 50 ° 20 ’ 47 ” W), 1 ♀, 05. I. 1985, A. A. Lise leg. (MCN 12739); Cambará do Sul, Itaim- bézinho (29 º 02 ’ 52 ” S; 50 º 08 ’ 41 ” W), 2 ♂, 1 ♀, 06. I. 1985, A. Lise leg. (MCN 12777; MCN 12779; MCN 12780); Garabi (28 ° 12 ’ 50 ” S; 55 ° 41 ’ 17 ” W), 1 ♀, XI. 1988, A. A. Lise leg. (IBSP 258334); Marcelino Ramos, Estreito Au- gusto Cesar (27 ° 24 ’ 51 ” S, 51 ° 27 ’ 2 ” W), 1 ♀, 03. II. 1990, C. A. Martinazzo leg. (MCN 19530); Maquiné (29 ° 40 ’ 30 ” S; 50 ° 12 ’ 25 ” W), Fepagro, 1 ♀, 18 – 27. I. 2002, Equipe Biota leg. (IBSP 139226); Derrubadas, Parque Estadual do Turvo (27 ° 8 ’ 44 ” S; 53 ° 53 ’ 10 ” W), 6 ♀ (IBSP 139987; IBSP 139989; IBSP 139990; IBSP 140007; IBSP 140018), all col- lected in 11 – 18. I. 2002 by Equipe Biota; Parque Estadual do Turvo (27 ° 8 ’ 44 ” S, 53 ° 53 ’ 10 ” W), 12 ♂, 3 ♀, 27 – 31. X. 2002 (MCN 37768); 1 ♂, 1 ♀, 19 – 22. X. 2004 (MCN 38812); 1 ♂, 27 – 31. X. 2003 (MCN 38813); 6 ♂, 27 – 31. X. 2003 (MCN 38814); 1 ♂, 19 – 22. X. 2004 (MCN 38836); 10 ♂, 6 ♀, 19 – 22. X. 2004 (MCN 38841), all collected by R. Ott et al.; Canela (29 ° 21 ’ 57 ” S; 50 ° 48 ’ 57 ” W), 1 ♀, 31. XII. 1973 (MCN 2070); 1 ♂, 20. III. 1976 (MCN 3854); 1 ♀, 05. III. 1977 (MCN 5209); 1 ♀, X. 1976 (MCN 6300), all collected by A. A. Lise; 1 ♂, 1 ♀, 03 – 04. IX. 1988, A. B. Bon- aldo leg. (MCN 17841); 1 ♀, 05. II. 1973 (MCN 852); 2 ♂, 31. XII. 1973 (MCN 2044); 1 ♀, 11. V. 1974 (MCN 2163); 1 ♂, 26. XII. 1974 (MCN 2475); 1 ♂, 1 ♀, 26. II. 1975 (MCN 3748); all collected by A. A. Lise; Carazinho (28 ° 17 ’ 2 ” S; 52 ° 47 ’ 9 ” W), 2 ♂, 10. XI. 1979, H. Bischoff leg. (MCN 8655); 1 ♀, 10. XI. 1979, A. A. Lise leg. (MCN 8686); So- bradinho (29 ° 25 ’ 15 ” S; 53 ° 1 ’ 44 ” W), 2 ♀, 10. I. 1985, A. A. Lise leg. (MCN 18717); Marcelino Ramos, Estreito Au- gusto Cesar (27 ° 24 ’ 51 ” S; 51 ° 27 ’ 2 ” W), 1 ♀, 03. II. 1990, C. A. Martinazzo leg. (MCN 19529); Nova Petrópolis, Boê- mios, Linha Imperial (29 ° 21 ’ 36 ” S; 51 ° 2 ’ 54.8 ” W), 1 ♂, 20. XII. 1973, A. A. Lise leg. (MCN 2206); Caxias do Sul, Vila Oliva (29 ° 12 ’ 56 ” S; 50 ° 53 ’ 22 ” W), 1 ♂, 10. I. 1976, P. A. Buckup leg. (MCN 3959); Caxias do Sul (29 ° 10 ’ 04 ” S; 51 ° 10 ’ 44 ” W), Água Azul, 1 ♀, 15. IX. 1976, H. Bischoff leg. (MCN 4532); 1 ♂, 04. IV. 1975, A. A. Lise leg. (MCN 18708); General Câmara (29 ° 54 ’ 18 ” S; 51 ° 45 ’ 36 ” W), 1 ♂, 1 ♀, 14. X. 1982, C. J. Becker leg. (MCN 10810); Far- roupilha (29 ° 13 ’ 30 ” S; 51 ° 20 ’ 52 ” W), 2 ♂, 1 ♀, 29. IX. 1978, H. Bischoff leg. (MCN 8283; MCN 8306; MCN 8329); Agudo, Usina Hidrelétrica Dona Francisca (29 ° 27 ’ 7 ” S; 53 ° 16 ’ 55 ” W), 1 ♀, 02. XI. 2002, A. B. B. Morais leg. (IBSP 122191); 1 ♀, Passo Fundo (28 ° 15 ’ 46 ” S 52 ° 24 ’ 25 ” W), 1 ♀, 13. X. 1985, A. A. Lise leg. (MCN 14321); 1 ♂, 01. VIII. 1986, A. A. Lise leg. (MCN 15573); Estrela Velha, Barragem de Itaúba (29 ° 15 ’ 40 ” S; 53 ° 14 ’ 8 ” W), 2 ♂, 1 ♀, 21. X. 1998, A. B. Bonaldo leg. (MCN 30606; MCN 30607); Garruchos (28 ° 11 ’ 2 ” S; 55 ° 38 ’ 20 ” W), 4 ♀, 10. XII. 1975 (MCN 3152); 3 ♀, 10. XII. 1975 (MCN 3172); 1 ♀, 06. XII. 1975 (MCN 18703); all collected by A. A. Lise; Triunfo (29 ° 56 ’ 34 ” S; 51 ° 43 ’ 4 ” W), 2 ♂, 1 ♀, 15. IX. 1977, H. Bischoff leg. (MCN 6516; MCN 6969); 1 ♂, 27. X. 1977, M. H. Galileo leg. (MCN 7031); 1 ♂, 2 ♀, 28. XI. 1977, A. A. Lise leg. (MCN 7393); Polo Petroquímico do Sul (29 ° 54 ’ 4 ” S; 51 ° 22 ’ 27 ” W), 1 ♂, 1 ♀, 20. X. 1977, E. H. Buckup leg. (MCN 18706; MCN 18707); Parobé (29 ° 37 ’ 44 ” S; 50 ° 50 ’ 6 ” W), Rio dos Sinos, 1 ♀, 10. XI. 2008, E. N. L. & P. E. S. Rodrigues leg. (MCN 47891); 1 ♀, 18. XII. 2007, E. N. L. & P. E. S. Rodrigues leg. (MCN 48029); Parque Braskem de Proteção Ambiental (29 ° 51 ’ 59.44 ” S; 51 ° 21 ’ 54.00 ” W), 2 ♂, 24. X. 1989, A. D. Brescovit leg. (MCN 18903); 1 ♂, 28. XI. 1989, E. H. Buckup leg. (MCN 19042); 1 ♀, 24. XI. 1999, M. A. L. Marques leg. (MCN 31739); 1 ♂, 21. X. 2003, A. Barcellos & L. Schmidt leg. (MCN 36458); 1 ♀, 2. XI. 2007, A. Barcellos & L. Schmidt leg. (MCN 43644); 1 ♀, 04. XII. 2009, M. R. M. Poeta leg. (MCN 46917); 2 ♂, 04. XII. 2009, M. Pairet Jr, leg. (MCN 46925); 1 ♀, 04. XII. 2009, P. E. S. Rodrigues leg. (MCN 46933); Arroio Grande, Mata Ciliar do Rio Piratini (31 ° 54 ’ 06.47 ” S; 52 ° 39 ’ 08.29 ” W), 1 ♀, 13. VI. 2008, E. N. L. & P. E. S. Rodrigues leg. (MCN 54732); Campo Bom (29 ° 40 ’ 44 ″ S; 51 ° 3 ’ 10 ” W), 1 ♀, 16. XI. 1979, C. J. Becker leg. (MCN 8789); Novo Hamburgo (29 º 40 ’ 40 ” S; 51 º 07 ’ 51 ” W), 2 ♂, 22. IX. 1986, C. J. Becker leg. (IBSP 221094); 1 ♂, 21. X. 1983 (MCN 11814); 1 ♀, 25. XI. 1983 (MCN 11847); 2 ♂, 11. XII. 1984 (MCN 12645); 2 ♂, 07. X. 1985 (MCN 14262; MCN 12646); 2 ♂, 14. XII. 1984 (MCN 12717); 1 ♀, 31. III. 1986, C. J. Becker leg. (MCN 14557), all collected by C. J. Becker; Morro Reuter (29 ° 32 ’ 16 ” S; 51 ° 4 ’ 51 ” W), 1 ♀, 28. III. 1976, C. J. Becker leg. (MCN 3576); Viamão, Estação Experimental Fitotécnica Águas Belas (30 ° 05 ’ 00 ” S; 51 ° 02 ’ 00 ” W), 1 ♂, 30. XI. 1984, M. A. L. Marques leg. (MCN 12611); 1 ♂, 06. I. 1977, A. A. Lise leg. (MCN 5899); São Leopoldo (29 º 45 ’ 36 ” S; 51 º 08 ’ 49 ” W), 1 ♂, 28. IX. 1987, C. J. Becker leg. (IBSP 221095); 2 ♂, 17. IX. 1982 (MCN 10965); 3 ♂, 24. IX. 1982 (MCN 11049); 2 ♂, 15. X. 1982 (MCN 11146); 1 ♀, 15. X. 1982 (MCN 11147); 1 ♀, 20. XII. 1982 (MCN 11339); 2 ♂, 12. XII. 1983 (MCN 11865); 1 ♂, 26. XII. 1983 (MCN 11911); all collected by C. J. Becker; Montenegro (29 ° 41 ’ 20 ” S; 51 ° 27 ’ 39 ” W), 2 ♂, 2 ♀, 06. X. 1977 (MCN 6806; MCN 7277); 3 ♂, 17. XI. 1977 (MCN 7278); 1 ♀, 20. XII. 1985 (MCN 14077); 4 ♂, 29. IX. 1977 (MCN 6718); 1 ♂, 17. XI. 1977, (MCN 7280); 8 ♂, 01. XII. 1977 (MCN 7482); 4 ♂, 20. XII. 1977 (MCN 7667); all collected by E. H. Buckup & A. A. Lise; Tapes (30 ° 40 ’ 22 ” S; 51 ° 23 ’ 45 ” W), Fazenda Guará, 1 ♀, 19. XII. 2003, Equipe Probio leg. (MCN 36821); Fazenda São Miguel, 2 ♂, 1 ♀, 17. XII. 2003, Equipe Probio leg. (MCN 36847); Barra do Ribeiro (30 ° 17 ’ 27 ” S; 51 ° 18 ’ 3 ” W), Fazenda Boa Vista, 2 ♂, 18. XII. 2003, Equipe Probio leg. (MCN 36850); Capão do Leão, Campus UFPEL (31 ° 45 ’ 46 ” S; 52 ° 29 ’ 2 ” W), 1 ♀, 24. II. 2000 (IBSP 36534); 2 ♂, 1 ♀, 30. IX. 2001 (IBSP 36536); 2 ♂, 30. IX. 2001 (IBSP 36537); 1 ♀, 30. IX. 2001 (IBSP 36538); all collected by E. N. L. Rodrigues. PARAGUAY. Departamento Paraguari: Parque Nacional de Ibycuí (26 º 05 ’ 49.08 ” S; 56 º 50 ’ 20.16 ” W), 1 ♀, 28. I. 1982 (MNHNP); 1 ♀, 20. X. 1981 (MNHNP); Departamento Amambay: Parque Nacional Cerro Corá, Base de Cerro Tuya Hog (22 ° 38 ’ 28 ” S; 56 ° 0 ’ 47 ” W), 1 ♀, X. 1983, (MNHNP); 1 ♀, 8. VI. 1982 (IBNP); Departamento Itapúa: Pirapó (28 ° 2 ’ 42 ” S; 55 ° 11 ’ 56 ” W), 1 ♂, 4. VIII. 1982 (MNHNP); Departamento Caaguazú: Caaguazú (25 ° 28 ’ 11 ” S; 56 ° 0 ’ 55 ” W), 1 ♂, 23. VII. 1982 (MNHNP); all collected by J. A. Kochalka. ARGENTINA. Misiones: Loreto (27 ° 19 ’ 55.12 ” S; 55 ° 31 ’ 23.47 ” W), 1 ♂, 27. III. 1932 – 8 / III / 1933, A. A. Oglobin leg. (MLP); Puerto Iguazú (25 ° 35 ’ 50 ” S; 54 ° 34 ’ 41 ” W), 8 ♀, XI. 1948, M. Birabén leg. (MLP); Puerto Victoria (26 ° 20 ’ 04 ” S; 54 ° 39 ’ 44 ” W), 1 ♀, [without date], A. Zenzes leg. (MLP); Parque Nacional del Iguazú, Cataratas del Iguazú (25 ° 41 ’ 43 ” S; 54 ° 26 ’ 12 ” W), 2 ♂, XI. 1989, (MACN- Ar); 1 ♀, XI. 1991, (MACN-Ar); 1 ♂, 1 ♀, XI. 1992, (MACN-Ar); all collected by M. J. Ramírez; Cantera, 200 m, 1 ♀, 8. XII. 1990 – 06. I. 1991, S. & J. Peck leg. (AMNH); Porto Canoas, 180 m, 1 ♀, 8. XII. 1990 – 06. I. 1992, S. & J. Peck leg. (AMNH); Eldorado (26 ° 28 ’ 00 ” S; 54 ° 43 ’ 00 ” W), 2 ♂, 01. X – 15. XI. 1964, A. Kovacs leg. (AMNH); 2 ♂, 1 ♀, 01. X – 15. XI. 1965, A. Kovacs leg. (AMNH).